From a networking perspective layer 7 of the OSI model is talking to a different layer which is not something network folks expect to see! This is why you must maintain 2007 Hub and CAS servers as long as you have 2007 mailboxes. This is due to the fact that RPC is used to communicate between the servers and is coded to only work with the relevant version. Conversely, the Exchange 2010 HUB Store Driver can only communicate with the Exchange 2010 Mailbox role. For example, the Exchange 2007 HUB Store Driver can only communicate with the Exchange 2007 Mailbox role. The intent was to remove the strict version dependency hooks which existed in the previous versions of Exchange. Exchange 2013 streamlined the connectivity between servers with the “every server is an island” design philosophy.
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